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Richard Gomulkiewicz

Richard Gomulkiewicz

Field of Study: Mathematical Population Biology and Evolution
Title: Professor
Degrees: Ph.D. University of California, Davis
Homepage: Homepage/Lab Web Site Link
Google Scholar:  Google Scholar
Office: Eastlick 391
Phone: 509-335-2527
Fax: nan
Mailing Address: School of Biological Sciences
Washington State University
PO Box 644236
Pullman,WA 99164-4236


I study theoretical population biology, population genetics, and evolutionary biology using mathematics. My research aim is to develop rigorous conceptual understanding of how complex evolutionary and demographic processes affect populations, species, and ecological communities. My investigations focus on how evolutionary and ecological mechanisms interact to determine the responses of organisms to each other and their environments including…
• The population-level and evolutionary responses of species facing novel or altered environments (“Evo Demo”).
• Coevolution in geographically structured populations and in multi-species communities (“Co-Evo Demo”).
• The evolution of environmentally sensitive phenotypes, a.k.a. phenotypic plasticity.
• The evolution of sexually selected traits and mating preferences.
• The evolution of “function-valued” traits.

I am also interested in applied biology, especially the development and interpretation of statistical tools based on molecular genetic data for assessing population characteristics of practical importance.

Representative Publications:

  • Aguiñaga, J., R. Gomulkiewicz, and H.E. Watts. (2021) Effect of social information on an individual’s assessment of its environment. Animal Behaviour. doi:
  • Gomulkiewicz, R., M.L. Thies, and J.J. Bull. (2021) Evading resistance to gene drives. Genetics 217 (2): iyaa040. doi:
  • Gomulkiewicz, R., J.G. Kingsolver, P.A. Carter, and N. Heckman. 2018. Variation and evolution of function-valued traits. Annual Review of Ecology, Evolution, and Systematics 49:139-64
  • Gomulkiewicz, R., S.M. Krone, and C.H. Remien. 2017. Evolution and the duration of a doomed population. Evolutionary Applications 10:471-484. doi: 10.1111/eva.12467
  • Drown, D. M., M. F. Dybdahl, and R. Gomulkiewicz. 2013. Consumer-resource interactions and the evolution of migration. Evolution 67:3290-3304. doi: 10.1111/evo.12194
  • Jones, E.I., S. L. Nuismer, and R. Gomulkiewicz. 2013. Revisiting Darwin’s conundrum reveals a twist on the relationship between phylogenetic distance and invasibility. Proceedings of the National Academy of Sciences 110 (51): 20627-20632. doi: 10.1073/pnas.1310247110
  • Barfield, M., R. D. Holt, and R. Gomulkiewicz. (2011) Evolution in stage-structured populations. The American Naturalist 177:397-409
  • Gomulkiewicz, R., R. D. Holt, M. Barfield, and S. L. Nuismer. (2010) Genetics, adaptation, and invasion in harsh environments. Evolutionary Applications 3:97-108
  • Gomulkiewicz, R., and D. Houle. (2009) Demographic and Genetic Constraints on Evolution. The American Naturalist 174:E218-E229
  • Gomulkiewicz, R., J. N. Thompson, R. D. Holt, S. L. Nuismer, and M. E. Hochberg. (2000.) Hot spots, cold spots, and the geographic mosaic theory of coevolution. The American Naturalist. 156:156-174
  • Gomulkiewicz, R., and R.D. Holt. (1995) When does evolution by natural selection prevent extinction? Evolution 49:201-207
  • Gomulkiewicz, R., and M. Kirkpatrick. (1992) Quantitative genetics and the evolution of reaction norms. Evolution 46:390-411