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Patrick Carter

Patrick Carter

Field of Study: Evolutionary Genetics and Evolutionary Physiology
Title: Professor
Degrees: PhD, Environmental, Population and Organismic Biology, University of Colorado, Boulder
Homepage: Homepage/Lab Web Site Link
Google Scholar:  Google Scholar
Office: Heald 217
Phone: 509-335-1447
Fax: nan
Mailing Address: School of Biological Sciences
Washington State University
PO Box 644236
Pullman,WA 99164-4236


My research focuses on the evolution of the integrated phenotype. Using ideas and methods from evolutionary and quantitative genetics, physiological ecology, and comparative physiology, my lab group addresses questions on the evolution of function-valued traits, the correlated responses of multiple traits to selection, and the relative effects of selection and drift on trait evolution. Our goal is to provide a fundamentally deeper understanding of the evolution of the entire phenotype and of evolutionary processes.

Representative Publications:

  • Bellinger, K, G.H. Thorgaard, and P.A. Carter. 2018. Complex relationship among rearing temperature, growth and sprint speed in clonal lines of rainbow trout. Canadian Journal of Fisheries & Aquatic Sciences 75:1868-1877.
  • Gomulkiewicz, R., J.G. Kingsolver, P.A. Carter, and N. Heckman. 2018. Variation and evolution of function-valued traits. Annual Review of Ecology, Evolution and Systematics 49:139-164.
  • Cabassi, A., D. Pigoli, P. Secchi, P.A. Carter. 2017. Permutation tests for the equality of covariance operators of functional data with applications to evolutionary biology. Electron. J. Statist. 11: 3815-3840. doi:10.1214/17-EJS1347.
  • Careau V., ME. Wolak, P.A. Carter & T. Garland Jr. 2015. Evolution of the additive genetic variance-covariance matrix under continuous directional selection on a complex behavioral phenotype. Proceedings of the Royal Society of London B 282:20151119.
  • Bellinger, K, G.H. Thorgaard, and P.A. Carter. 2014. Effects of domestication on growth and swim performance in rainbow trout. Aquaculture 420-421:154-159.
  • Gervini, D. and P.A. Carter. 2014. Warped functional analysis of variance. Biometrics 70:526-353.
  • Irwin, K.K., and P.A. Carter. 2014. Artificial selection on larval growth curves in Tribolium castaneum: Correlated responses and constraints. Journal of Evolutionary Biology 27:2069-2079.
  • Careau, V., M.E. Wolak, P.A. Carter, and T. Garland, Jr. 2013. Limits to behavioral evolution: the quantitative genetics of a complex trait under directional selection. Evolution 67:3102-3119.
  • Heink, A.E., A. Parrish, G.H. Thorgaard and P.A. Carter. 2013. Oxidative stress among SOD-1 genotypes in rainbow trout (Oncorhynchus mykiss). Aquatic Toxicology 144-145: 75-82.
  • Irwin, K.K., and P.A. Carter. 2013. Constraints on the evolution of function-valued traits: A study of growth in Tribolium castaneum. Journal of Evolutionary Biology 26:2633-2643.