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Steve Bollens

Steve Bollens

Field of Study: Biological Sciences
Title: Professor and Director Science Programs (Vancouver)
Degrees: Ph.D., Biological Oceanography, University of Washington
Homepage: Homepage/Lab Web Site Link
Google Scholar:  Google Scholar
Office: Science (VSCI) 230Q
Phone: 360-546-9116
Mailing Address: School of Biological Sciences
Washington State University
PO Box 644236
Pullman,WA 99164-4236


My research is broadly concerned with the ecology of marine and estuarine zooplankton and fish, and spans the sub-disciplines of behavior, population biology, community ecology and ecosystem dynamics. My research often has an applied aspect to it, touching upon such areas as conservation biology, restoration ecology, fisheries oceanography, and global change. I employ a wide variety of approaches to “doing science”, including field (observational), modeling and experimental techniques. Locations of my current or recent projects include estuaries of the northeast Pacific (CA, OR, WA, and BC), Georges Bank/Northwest Atlantic, the Arabian Sea, the Florida Keys, and the Bering Sea. These projects are funded by a wide range of federal and state agencies, including the National Science Foundation (NSF), the Environmental Protection Agency (EPA), the Office of Naval Research (ONR), the National Oceanic and Atmospheric Administration (NOAA), the United States Geological Survey (USGS), and the CALFED Bay/Delta Program.

Representative Publications:

  • Bollens, S. M., J. C. Cordell, S. Avent, and R. Hooff. 2002. Invasive Zooplankton: A Brief Review, Plus Two Case Studies from the Northeast Pacific Ocean. Hydrobiologia, 480: 87-110.
  • Lougee, L., S. M. Bollens, and S. R. Avent. 2002. The effect of haloclines on vertical distribution and migration of zooplankton. J. Exp. Mar. Biol. Ecol. 278: 111-134.
  • Beet, A., A. R. Solow, and S. M. Bollens. 2003. Comparing vertical plankton profiles with replication. Mar. Ecol. Prog. Ser., 262:285-287.
  • Purkerson, D. G., M. A. Doblin, S. M. Bollens, S. N. Luoma, G. A. Cutter. 2003. Selenium in San Francisco Bay Zooplankton: Potential Effects of Hydrodynamics and Food Web Interactions. Estuaries. 26: 956-969.
  • Bochdansky, A. B. and S. M. Bollens. 2004. Relevant scales in zooplankton ecology: Distribution, feeding and reproduction of the copepod Acartia hudsonica in response to thin-layers of the diatom Skeletonema costatum. Limnol. Oceanogr. 49: 625-636.
  • Bollens, S. M., and A. Sanders. 2004. Ecology of larval Pacific herring Clupea pallasi in the San Francisco Estuary: Seasonal and interannual abundance, distribution, diet and condition. Amer. Fish. Soc. Symp. 36: 15-35.
  • Brown, H., S. M. Bollens, L. P. Madin, and E. F. Horgan. 2004. Effects of Warm Water Intrusions on Populations of Macrozooplankton and Micronekton on Georges Bank, Northwest Atlantic. Cont. Shelf Res. In Press.
  • Clay, T. W, S. M. Bollens, A. B. Bochdansky, and T. R. Ignoffo. 2004. The Effects of Thin Layers on the Vertical Distribution of Larval Pacific Herring, Clupea pallasi. J. Exp. Mar. Biol. Ecol. 305: 171-189.
  • Dorman, J.G., S. M. Bollens and A. M. Slaughter. 2004. Population Biology of Euphausiids off Northern California, with Special Reference to the Effects of Event-Scale Winds on Euphausia pacifica. Mar. Ecol. Prog. Ser. In Press.
  • Gewant, D. S. and S. M. Bollens. 2004. Macrozooplankton and Micronekton of the Lower San Francisco Estuary: Seasonal, Interannual and Regional Variation in Relation to Environmental Conditions. Estuaries, Accepted pending revisions.
  • Hooff, R. C. and S. M. Bollens. 2004. Functional response and potential predatory impact of Tortanus dextrilobatus, a recently introduced carnivorous copepod in the San Francisco Estuary. Mar. Ecol. Prog. Ser. 277: 167-179.
  • Visintainer, T. A., S. M. Bollens, and C. Simenstad. 2004. Community Composition and Diet o of Fishes as a Function of Tidal Channel Order: A Field Study in China Camp Marsh, San Francisco Estuary. Mar. Ecol. Prog. Ser., Accepted pending revisions.
  • Dean, A. F., S. M. Bollens, C. Simenstad, and J. Cordell. 2004. Population Dynamics of the Mysid Neomysis Kadiakensis and Flux Between a Tidal Marsh and Open Water in The San Francisco Estuary. Estuar. Coast. Shelf Sci., Accepted pending revisions.
  • Ignoffo, T., S. M. Bollens, and A. Bochdansky. 2004. The Effect of Thin Layers on the Vertical Distribution of the Rotifer Brachionus plicatilis. J. Exp. Mar. Biol. Ecol. In Press.